(1985), in referring to the preliminary stages of courtship in his diagrams (Halliday 1990). It was characteristic of Tim that he delighted in using the term “spermatophoreplay”, coined by Houck et al. On top of that, Tim’s papers were illustrated with beautiful diagrams and portraits of displaying newts. His approach was observational, quantitative, comparative, and experimental – a tour de force that must have impressed Niko and the rest of the Group. Throughout this period (1968-1976) Tim published ground-breaking, synthetic work on newt courtship. Cullen for his DPhil and then carried on in the ABRG as a post-doc. Tim joined the Animal Behaviour Research Group (ABRG) in the Zoology department, led by Niko Tinbergen, in 1968 and began his studies of courtship behaviour in newts. After graduating he trained to be a teacher at Cambridge but then decided he would rather do a DPhil and returned to Oxford. Fascinated by newts and talented at drawing as a child, Tim decided to study Zoology at Oxford University. Jack was a housemaster and taught biology at Marlborough College in Wiltshire, England, whilst Edna was a housemistress. Tim was the youngest son of Jack and Edna Halliday. Tim Halliday was one of the fathers of the study of sexual selection and a leading light in amphibian conservation, he was also a talented illustrator and will be sorely missed by his family, friends and colleagues. Timothy Richard Halliday, 73, a zoologist, conservationist and artist, died on 10 th April 2019, after a long illness caused by lymphoma, in Oxford, England.
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